All organisms are communities of interacting coevolving replicators of different classes. Three other organisms within this genus are causes of nonvenereal or endemic treponematoses.
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Pallidum subspecies pertenue is the causative agent of yaws T.
. Pallidum subspecies endemicum causes endemic non-venereal syphilis and T. These pathogens are morphologically and antigenically indistinguishable. A complete theory of replicator coevolution remains to be developed but it appears likely that not only the differentiation between selfish and cooperative replicators but the emergence of the entire range of replication strategies from selfish to cooperative is intrinsic to biological.
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